This article was downloaded by: [Nicholas Famoso] On: 18 September 2014, At: 11:35 Publisher: Taylor & Francis Informa Ltd Registered in England and Wales Registered Number: 1072954 Registered office: Mortimer House, 37-41 Mortimer Street, London W1T 3JH, UK Journal of Vertebrate Paleontology Publication details, including instructions for authors and subscription information: http://www.tandfonline.com/loi/ujvp20 Correction to the holotype (AMNH FM 9394) of Merychippus proparvulus Osborn, 1918 (Perissodactyla, Equidae) Nicholas A. Famosoa & Samantha S. B. Hopkinsb a Department of Geological Sciences and Museum of Natural and Cultural History, University of Oregon, 1272 University of Oregon, Eugene, Oregon 97403, U.S.A. b Clark Honors College and Department of Geological Sciences, University of Oregon, 1272 University of Oregon, Eugene, Oregon 97403, U.S.A. Published online: 09 Sep 2014. To cite this article: Nicholas A. Famoso & Samantha S. B. Hopkins (2014) Correction to the holotype (AMNH FM 9394) of Merychippus proparvulus Osborn, 1918 (Perissodactyla, Equidae), Journal of Vertebrate Paleontology, 34:5, 1249-1250, DOI: 10.1080/02724634.2014.853073 To link to this article: http://dx.doi.org/10.1080/02724634.2014.853073 PLEASE SCROLL DOWN FOR ARTICLE Taylor & Francis makes every effort to ensure the accuracy of all the information (the “Content”) contained in the publications on our platform. However, Taylor & Francis, our agents, and our licensors make no representations or warranties whatsoever as to the accuracy, completeness, or suitability for any purpose of the Content. Any opinions and views expressed in this publication are the opinions and views of the authors, and are not the views of or endorsed by Taylor & Francis. The accuracy of the Content should not be relied upon and should be independently verified with primary sources of information. 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Terms & Conditions of access and use can be found at http:// www.tandfonline.com/page/terms-and-conditions http://www.tandfonline.com/loi/ujvp20 http://www.tandfonline.com/action/showCitFormats?doi=10.1080/02724634.2014.853073 http://dx.doi.org/10.1080/02724634.2014.853073 http://www.tandfonline.com/page/terms-and-conditions http://www.tandfonline.com/page/terms-and-conditions SHORT COMMUNICATION CORRECTION TO THE HOLOTYPE (AMNH FM 9394) OFMERYCHIPPUS PROPARVULUS OSBORN, 1918 (PERISSODACTYLA, EQUIDAE) NICHOLAS A. FAMOSO*,1 and SAMANTHA S. B. HOPKINS2; 1Department of Geological Sciences and Museum of Natural and Cultural History, University of Oregon, 1272 University of Oregon, Eugene, Oregon 97403, U.S.A., nfamoso2@uoregon.edu; 2Clark Honors College and Department of Geological Sciences, University of Oregon, 1272 University of Oregon, Eugene, Oregon 97403, U.S.A., shopkins@uoregon.edu On a recent research trip to the American Museum of Natural History (AMNH), it was discovered that the holotype of Mery- chippus proparvulus Osborn, 1918:117 (Perissodactyla, Equi- dae), included postcrania from a camelid. Camelid postcrania originally cataloged with the M. proparvulus holotype include a distal metapodial, right calcaneum, and proximal phalanx (Fig. 1). The holotype specimen (upper left toothrow, cervical, thoracic, and caudal vertebrae, left hind foot [with digits II, III, and IV], and parts of limb bones [Osborn, 1918:pls. 11.3, 15.6, and 54.1, Fig. 91]) and presumably the associated camelid post- crania are from the Pawnee Creek beds, Pawnee Buttes, of northeastern Colorado (Osborn, 1918). Pawnee Creek is a classic late Barstovian (Miocene) site, dated at 14.5–14 Ma (Tedford, 2004; Tedford et al., 2004). The three elements have morphology that clearly designates them as not only artiodactyls, but more specifically camelid. The distal metapodial is clearly a fusion of metapodials III and IV over the proximal end and most of the shaft (diagnostic of derived artiodactyls), but the two remain unfused distally, a fea- ture unique to camels and putatively related to their adaptation to a pacing gait (Peterson, 1904; Breyer, 1983; Janis et al., 2002). The calcaneum has wide cuboid facets, the tuber calcis of the FIGURE 1. Camelid specimens labeled with the same permanent number as the type of Merychippus proparvulus, AMNH 9394. A, proximal phalanx; B, calcaneum; C, distal metapodial. Scale bar equals 5 cm. *Corresponding author. 1249 Journal of Vertebrate Paleontology 34(5):1249–1250, September 2014 � 2014 by the Society of Vertebrate Paleontology D ow nl oa de d by [ N ic ho la s Fa m os o] a t 1 1: 35 1 8 Se pt em be r 20 14 calcaneum has a blunt cap, and the corpus calcanei is elongate, all diagnostic features of camelid calcanea (Peterson, 1904; Webb, 1965). The proximal phalanx is elongate and slender with a distal articular surface that extends dorsally onto the volar sur- face, illustrative of the semidigitigrade posture adopted by cam- els (Peterson, 1904; Janis et al., 2002). The carinal groove on the posterior border of the proximal articular surface is wide and shallow, also a diagnostic feature of camelid postcrania (Peter- son, 1904; Webb, 1965; Breyer, 1983; Janis et al., 2002). The American Museum of Natural History has assigned new num- bers to the camelid specimens; AMNH FM 144426 to the pha- lanx, AMNH FM 144427 to the calcanium, and AMNH FM 144428 to the metapodial. The original number, AMNH FM 9394, now contains an upper left toothrow, cervical, thoracic, and caudal vertebrae, left hind foot (with digits II, III, and IV), and parts of limb bones, of which only the foot and toothrow were figured by Osborn (1918) and other elements that appear to be diagnosable to Equidae, including a row of lower incisors with an associated canine, a partial right upper toothrow, left radio-ulna, one II phalanx (digit III), one III phalanx (digit III), and an articulated phalanx II and III from digit III. The remain- der of the elements (three ribs, several rib fragments, and bone fragments) in this specimen number cannot be assigned with con- fidence to either Perissodactyla or Artiodactyla, and so we refrain from separating these elements from the original specimen number. ACKNOWLEDGMENTS We would like to thank J. Meng, R. O’Leary, and A. Gishlick for recataloging the camelid material at the AMNH, M. Emery for photographing the equid material for our reference, and M. Woodburne and P. Holroyd for constructive comments. N. A. F. was supported by a Geological Society of America Graduate Student Research Grant and the Paleontological Soci- ety Richard K. Bambach Award on this particular research trip. LITERATURE CITED Breyer, J. A. 1983. The biostratigraphic utility of camel metapodials. Journal of Paleontology 57:302–307. Janis, C. M., J. M. Theodor, and B. Boisvert. 2002. 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Woodburne (ed.), Late Cretaceous and Cenozoic Mammals of North America. Columbia University Press, New York. Webb, S. D. 1965. The Osteology of Camelops. Bulletin of the Los Angeles County Museum Science: Number 1. Submitted July 10, 2013; revisions received September 25, 2013; accepted September 30, 2013. Handling editor: Thomas Martin. 1250 JOURNAL OF VERTEBRATE PALEONTOLOGY, VOL. 34, NO. 5, 2014 D ow nl oa de d by [ N ic ho la s Fa m os o] a t 1 1: 35 1 8 Se pt em be r 20 14