TYPE SPECIMENS 
AT THE UNIVERSITY OF OREGON: 
FOSSIL VERTEBRATES AND PLANTS 
by 
ERIC PAUL GUSTAFSON and JOYCE EATON 
Bulletin No. 24 
Museum of Natural History 
University of Oregon 
Eugene, Oregon 
Marchl983 
The Bulletin of the Museum of Natural History of the University of Oregon is published to 
increase the knowledge and understanding of the natural history of Oregon. Original articles, 
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Cover-Oreodon superbus Leidy, as shown in Plate 1, Figure 1, of Joseph Leidy, Contributions 
to the Extinct Vertebrate Fauna of the Western Territories, 1873 ( see Literature Cited). 
Volume editor-Don E. Dumond 
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Notes on Some Upper Miocene Shrews from Oregon, by J. H. Hutchison; 23 pages, 17 
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Peromyscus of the Late Tertiary in Oregon, by J. Arnold Shotwell; 35 pages, 11 figures, 
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[Continued inside back cover. l 
TYPE SPECIMENS 
AT THE UNIVERSITY OF OREGON: 
FOSSIL VERTEBRATES AND PLANTS 

TYPE SPECIMENS 
AT THE UNIVERSITY OF OREGON: 
FOSSIL VERTEBRATES AND PLANTS 
by 
Emc PAUL GusTAFSON and ]oYCE EATON 
Bulletin No. 24 
Museum of Natural History 
University of Oregon 
Eugene, Oregon 
Marchl983 

I 
1 
Type Specimens at the University of Oregon: 
Fossil Vertebrates and Plants 
by 
ERIC PAUL GUSTAFSON and JoYCE EATON 
INTRODUCTION 
A published list of type specimens such as this 
serves several purposes. First and foremost it is in-
tended to allow quick access to the type holdings 
of a museum and to the basic information that is 
present in the museum files. Secondly, it allows some 
evaluation of the scientific value of a given collection. 
This latter can be particularly significant in the case 
of a collection such as that at the University of Ore-
gon, which is of considerable size ( 13th in the United 
States in terms of total catalogued specimens of fossil 
vertebrates as of 1977, according to Langston et al. , 
1978) but is off the beaten paths which in Vertebrate 
Paleontology lead back and forth between California 
and the northeastern states. We hope that this publi-
cation will also serve as a biblographic introduction 
to the widespread literature on fossil vertebrates from 
Oregon. It certainly will not serve as a complete guide 
to fossil vertebrate type specimens from the State of 
Oregon ; these are widespread in other collections 
and only a small fraction of the taxonomic work on 
the various fossil faunas had been done here. 
The large collection of fossil plant materials at the 
University of Oregon, mostly leaf impressions, is 
poorly documented in the literature. Thus, there is 
relatively little to report in the way of type specimens. 
Much of the bibliographic work involved in this 
publication was done by Joyce Eaton in early 1978 
while helping to reexamine and improve the curation 
of the UO type collections. The organization and 
editing of this list are entirely the responsibility of 
the senior author. 
HrsTORY oF THE CoLLECTIONs 
At the date of this publication, the specimens here-
in listed are housed in the University of Oregon 
Museum of Natural History building, but are a sepa-
rate administrative responsibility under the title of 
Condon Museum of Geology in the Department of 
Geology. The history of these collections has some 
complications. Originally the "University of Oregon's 
collection" was, in fact, the private collection of Dr. 
Thomas Condon, one of the first members of the 
faculty of the University when it was organized in 
1876. He had begun his collecting activities in the 
1840's, but his collection of vertebrate fossils was 
begun while he was a minister at The Dalles, Oregon, 
in the 1860's. When Dr. Condon died in 1907 his 
large collection remained with the University, and 
when purchased in 1909 from his heirs, it became the 
Condon Museum of Geology. In 1936 the Condon 
Collection ( with some additions) was added to the 
newly organized University of Oregon Museum of 
Natural History. The bulk of the present collection 
was added in the years 1946-1971 when Dr. J. Arnold 
Shotwell was Curator of the collections and (for 
some of those years) Director of the museum. In 
September of 1977 the old Condon Museum collec-
tion and other specimens related to geology and pale-
ontology were placed in the care of the Department 
of Geology as part of a general reorganization of the 
museums. 
CONTENT OF THIS CATALOG 
Only those specimens listed in the original publi-
cations as types, holotypes, cotypes, or paratypes, are 
listed here. The Condon Museum collections contain 
between 800 and 1000 hypotypes ( described, figured, 
or listed specimens) which could not be included. 
Entries to the list follow this pattern: The specific 
name is listed in boldface type as originally pub-
lished, followed by the most recent revision ( if any) 
in italics. Then follows the type status, the specimen 
number and description, the locality number and 
description as originally given, the stratigraphic po-
sition, the age ( as originally given and as revised), 
the collector, and any comments that seem necessary 
or desirable for understanding the history of the 
specimen. 
2 BULLETIN, MUSEUM OF NATURAL HISTORY, UNIVERSITY OF OREGON No.24 
VERTEBRATES 
Phylum CHORDATA 
Subphylum VERTEBRATA 
Class OSTEICHTHYES 
Order SILURIFORMES 
Family Ictaluridae 
* Ictalurus peregrinus Lundberg, 1975 
Type status-Holotype 
Specimen-F-28035, left pectoral spine 28.3 mm in 
length. 
Locahty-UO 2337, Sec. 11 (14 in original pub.), 
T21S, R37E, WM, Black Butte local fauna. 
Stratigraphic position- upper member of Juntura 
Formation, Malheur County, Oregon. 
Age-Clarendonian, Lower Pliocene (Upper 
Miocene). 
Collector-Shotwell. 
Order SALMONIFORMES 
Family Salmonidae 
*Smilodonichthys rastrosus Cavender and Miller, 
1972 
Type status-Holotype 
Specimen-F-26799, skull with jaws and anterior 
gill arches. 
Locality-VO 2250, gravel quarry just southwest 
of Gateway, Madras Quadrangle, Jefferson County, 
Oregon. 
Stratigraphic position-Madras Formation (also 
called Deschutes Fm. in some publications), from a 
sand lens within the "torrential gravels" in the lower 
part of the quarry. 
Collector-liames and McKain, 1964. 
* Smilodonichthys rastrosus Cavender and Miller, 
1972 
Type status-Paratype 
Specimen-F-3335, associated bones from a single 
individual, including crushed neurocranium, left and 
right mandibles and premaxillae, about 40 vertebrae, 
etc. 
Locality-VO 2250, same as holotype. 
Stratigraphic position-same as holotype. 
Collector-}. A. Shotwell, 1950. 
Class AMPHIBIA 
Order CAUDATA 
Family Salamandridae 
* Paleotaricha oligocenica van Frank, 1955 
Type status-Holotype 
Specimen-F-5405, complete skeleton on slab. 
Locality-VO 2715, Plant-bearing tuffs (locality 
of "Willamette Flora") 3 miles southeast of Eugene, 
/'I mile north of Goshen, Lane County, Oregon. 
Stratigraphic position-Tuffaceous shales and 
sandstones apparently overlying the Fisher and 
Eugene Formations. 
Age-Upper Oligocene. 
Collector-George Moorhead, 1951. 
Comments-The exposed surface of this specimen 
has been completely covered with transparent plastic 
to allow preparation of the underside of the skull. 
Class AVES 
Order GALLIFORMES 
Family Phasianidae 
* Lophortyx shotwelli Brodkorb, 1958 
Type status-Holotype 
Specimen-F -3611, prox1.mal portion of left 
humerus. 
Locality-VO 2222, East bank of McKay 
Reservoir, Pendleton Quadrangle, Umatilla County, 
Oregon. 
Stratigraphic position-Tuffaceous sandstone 
filling a channel cut into upper flows of the Yakima 
Basalt. 
Age-Pliocene, Hemphillian (probably very late 
Hemphillian) . 
Collector-]. A. Shotwell and Genevieve Shotwell, 
1950. 
Order CHARADRIFORMES 
Family Scolopacidae 
*Bartramia umatilla Brodkorb, 1958 
Type status-Holotype 
Specimen-F-3727, right carpometacarpus. 
Locality-VO 2222, east bank of McKay Reservoir, 
Pendleton Quadrangle, Umatilla County, Oregon. 
Stratigraphic position-Tuffaceous sandstone in 
channel cut into Yakima Basalt. 
Age-Pliocene, Hemphillian (probably very late 
Hemphillian). 
Collector-}. A. Shotwell and Genevieve Shotwell, 
1950. 
Order PELECANIFORMES 
Family Phalacrocoracidae 
*Phalacrocorax leptopus Brodkorb, 1961 
Type status-Holotype 
1983 GUST AF SON AND EATON: TYPE SPECIMENS 3 
Specimen-F-7994, proximal half of left tarso-
metatarsus. 
Locality-VO 2360, Malheur County, Oregon. 
Stratigraphic position- Upper member of 
Juntura Formation. 
Age-Lower Pliocene, Clarendonian (Upper 
Miocene). 
Collector- Shotwell and Russell, 1955. 
Order ARDEIFORMES 
Family Palaelodidae 
*Megapaloelodus opsigonus Brodkorb, 1961 
Type status-Holotype 
Specimen-F-5459, proximal end of left tarso-
metatarsus. 
Locality- VO 2334, SW:!4, Sec. 7, T21S, R38E, 
WM., Malheur County, Oregon. 
Stratigraphic position-Upper member of J untura 
Formation. 
Age-Lower Pliocene, Clarendonian (Upper 
Miocene). 
Collector-Shotwell and Alvey, 1954. 
Order ANSERIFORMES 
Family Anatidae 
*Eremochen russelli Brodkorb, 1961 
Type status-Holotype 
Specimen- F-5414 and F-5424, proximal part of 
humerus. 
Locality-VO 2335, SW_y,i, Sec. 7, T21S, R38E, 
WM, Malheur County, Oregon. 
Stratigraphic position-Upper member of Juntura 
Formation. 
Age-Lower Pliocene, Clarendonian (Upper 
Miocene). 
Collector- Shotwell and Alvey, 1954. 
*Eremochen russelli Brodkorb, 1961 
Type status-Paratype 
Specimen- F-5872 , coracoid. 
Locality-VO 2338, W Sec. 11, T21S, R37E, WM, 
Malheur County, Oregon. 
Stratigraphic position-} untura Formation, upper 
member, 50 feet below pumice band. 
Age- Lower Pliocene, Clarendonian (Upper 
Miocene). 
Collector-Shotwell and Alvey, 1954. 
*Querquedula pullulans Brodkorb, 1961 
Type status-Holotype 
Specimen-F-6289, proximal part of left carpo-
metacarpus. 
Locality-VO 2337, Sec. 11, T21S, R37E, Malheur 
County, Oregon. 
Stratigraphic position-Upper member of Juntura 
Formation. 
Age- Lower Pliocene, Clarendonian (Upper 
Miocene). 
Collector-Shotwell, 1954. 
* Ocyplonessa shotwelli Brodkorb, 1961 
Type Status-Holotype 
Specimen-F-11291 and F-10485, distal part of 
left tarsometarsus. 
Locality-VO 2337, Sec. 11, T21S R37E, WM, 
Malheur County, Oregon. 
Stratigraphic position- Upper member of Juntura 
Formation. 
Age- Lower Pliocene, Clarendonian (Upper 
Miocene). 
Collector-Shotwell, 1954. 
Order RALLIFORMES 
Family Rallidae 
*Fulica intelix Brodkorb, 1961 
Type status- Holotype 
Specimen-F-5758, distal portion of left 
tibiotarsus. 
Locality-VO 2341, Malheur County, Oregon. 
Stratigraphic position- Upper member of Juntura 
Formation, immediately below the pumice band. 
Age-Lower Pliocene, Clarendonian (Upper 
Miocene). 
Collector-Shotwell and Alvey, 1954. 
Class MAMMALIA 
Order INSECTIVORA 
Family Soricidae 
* Alluvisorex arcadentes Hutchison, 1966 
Type status-Holotype 
Specimen-F-22307, left dentary with 13-M2. 
Locality-VO 2465, west side of Quartz Basin, 
center of Sec. 33, T24S, R43E, WM, Malheur County, 
Oregon. 
Stratigraphic position-Deer Butte Formation. 
Age-Miocene, Barstovian. 
Collector-Shotwell, 1960. 
*Trimylus mawbyi Repenning, 1967 
Type status-Holotype 
Specimen-F-10486, right mandible with I, P 1, 
P4, M1-H· 
Locality-5 miles southwest of south end of Guano 
Lake, and about 25 miles southwest of Beatty Butte, 
Lake County, Oregon. 
Stratigraphic position-not known. 
Age- Miocene, Barstovian. 
Collector-F. B. Van Houten, about 1966. 
4 BULLETIN, MUSEUM OF NATURAL HISTORY, UNIVERSITY OF OREGON No.24 
* Paradomnina relictus Hutchison, 1966 
Type status- Holotype 
Specimen- F-24279, left mandible with P~-M:1. 
Locality- UO 2495, Skull Springs, SE corner Sec. 
10, T23S, R40E, WM, Malheur County, Oregon. 
Stratigraphic position-Butte Creek Volanic 
Sandstone. 
Age- Miocene, Barstovian. 
Collector- Shotwell , 1961. 
*Ingentisorex tumididens Hutchison, 1966 
Type status- Holotype 
Specimen- F-21960, left mandible with P, -M2. 
Locality-UO 2465, west side of Quartz Basin, cen-
ter of Sec. 33, T24S, R43E, WM, Malheur County, 
Oregon. 
Stratigraphic position-Deer Butte Formation. 
Age- Miocene, Barstovian. 
Collector- Shotwell, 1960. 
* Mystipterus (Mystipterus) pacificus Hutchison 
1968 
Type status- Holotype 
Specimen-UO F-22438, incomplete left dentary 
with P 4-M~, roots of P ~ and posterior half of P 2· 
Locality-UO 2465, Quartz Basin, west side, center 
of Sec. 33, T24S, R43E, WM, Malheur County, 
Oregon. 
Stratigraphic position-Deer Butte Formation. 
Age-Miocene (Barstovian). 
Collector-Shotwell, 1960. 
*Scalopoides ripafodiator Hutchison 1968 
Type status-Holotype 
Specimen- F-22488, right manidle with 1z and 
Pi-Ma. 
Locality-UO 2465, Quartz Basin, west side, center 
of Sec. 33, T24S, R43E, Malheur County Oregon. 
Stratigraphic position-Deer Butte Formation. 
Age-Miocene (Barstovian). 
Collector-Shotwell, 1960. 
*Scapanus (Xeroscapheus) proceridens 
Hutchison 1968 
Type status-Holotype 
Specimen-lJO F-22503, right mandible with 12, 
P 1,M1- ~. 
Locality-UO 2323, Krebs Ranch II; NW14 Sec. 
31, T3N, R22E, WM, Arlington Quad., Gilliam 
County, Oregon. 
Stratigraphic position-Dalles or Ellensburg 
Formation, above Elephant Mountain basalt flow. 
Age-Pliocene ( Hemphillian; possibly late 
Miocene). 
Collector-Shotwell, 1954. 
* Scapanoscapter simplicidens Hutchison 1968 
Type status-Holotype 
Specimen-UO F-24286, left mandible with P1, 
partial M1, M2-a-
Locality- UO 2495, Red Basin, Skull Springs 
fauna , SE corner Section 10, T23S, R40E, Malheur 
County, Oregon. 
Stratigraphic position-Butte Creek Volcanic 
Sandstone. 
Age-Miocene (Barstovian). 
Collector-Shotwell, 1961. 
* Achlyoscapter longirostris Hutchison, 1968 
Type status-Holotype 
Specimen-UO F-22412, left mandible with Ia-M1. 
Locality-UO 2465, Quartz Basin, west side, center 
of Sec. 33, T24S, R43E, WM, Malheur County, 
Oregon. 
Stratigraphic position-Deer Butte Formation. 
Age-Miocene (Barstovian). 
Collector-Shotwell, 1960. 
* ?Neurotrichus columbianus Hutchison 1968 
Type status-Holotype 
Specimen-UO F-24816, nearly complete right 
mandible. 
Locality-UO 2222, McKay Reservoir, east bank, 
Pehdleton Quad., Umatilla County, Oregon. 
Stratigraphic position-Tuffaceous sandstone in 
channel cut into upper Yakima Basalt. 
Age-Pliocene ( Hemphillian) . 
Collector-Shotwell, 1949. 
*Hydroscapheus americanus Shotwell, 1956 
Type status-Holotype 
Specimen-F-4101, humerus. 
Locality-UO 2222, McKay Reservoir, Pendleton 
Quad., Umatilla County, Oregon. 
Stratigraphic position-Tuffaceous sandstones in 
channel cut into upper Yakima Basalt. 
Age-Pliocene (Hemphillian). 
Collector-Shotwell, 1949. 
*Hydroscapheus americanus Shotwell, 1956 
Type status-Paratypes 
Specimen-F-3206, clavicle; F-2698 femur; F-
2456 and F-3792, tibia-fibulae. 
Locality-UO 2222, same as holotype. 
Age-Pliocene (Hemphillian). 
Collector-Shotwell, 1949. 
Order RODENTIA 
*Sewelleladon predontia Shotwell 1958 
Type status-Holotype 
1983 GUST AF SON AND EATON: TYPE SPECIMENS 5 
Specimen- F-4 734, left mandible with P 4-M3. 
Locality-VO 2275, along Haystack Creek, 3 miles 
NE of Spray, Wheeler County, Oregon. 
Stratigraphic position-John Day Formation 
( lower part of upper member) , from a slump block. 
Age-Miocene (Arikareean). 
Collector-Shotwell et al. 1946. 
*Citellus (Otospermophilus) wilsoni Shotwell, 
1956 
Type status- Parat ype 
Specimen- F-3596 , crushed skull and lower jaws, 
fore and hind feet, radii and ulna, distal tibia. 
Locality- VO 2222, McKay Reservoir, east bank, 
Pendleton Quad. , Umatilla, Oregon. 
Stratigraphic position-Tuffaceous sandstone in 
channel cut into Yakima Basalt. 
Age-Pliocene ( Hemphillian) . 
Collector- Shotwell, 1949. 
*Marmota oregonensis Shotwell , 1956 
Type status- Holot ype 
Specimen-F-3625, left M1 • 
Locality-VO 2222, McKay Reservoir, east bank, 
Pendleton Quad. , Umatilla, County, Oregon. 
Stratigraphic position-Tuffaceous sandstone in 
channel cut into Yakima Basalt. 
Age-Pliocene (Hemphillian). 
Collector-Shotwell, 1949. 
*Marmota oregonensis Shotwell, 1956 
Type status-Paratype 
Specimen-F-3626, right M1. 
Locality-VO 2222, McKay Reservoir, east bank, 
Pendleton Quad., Umatilla County, Oregon. 
Stratigraphic position-Same as holotype. 
Age-Pliocene (Hemphillian). 
Collector-Shotwell, 1949. 
*Citellus (Citellus) mckayensis Shotwell, 1956 
Type status-Holotype 
Specimen- F-3621, left mandible fragment with 
M1-a, 
Locality- VO 2222, McKay Reservoir, east bank, 
Pendleton Quad., Umatilla County, Oregon. 
Stratigraphic position-Tuffaceous sandstone in 
channel cut into Yakima Basalt. 
Age-Pliocene (Hemphillian). 
Collector- Shotwell, 1949. 
*Citellus (Otospermophilus) wilsoni Shotwell, 
1956 
Type status-Holotype 
Specimen-F-4091, right lower jaw with P4-M3. 
Locality-McKay Reservoir, east bank. Pendleton 
Quad., Umatilla County, Oregon. 
Stratigraphic position-Tuffaceous sandstone in 
channel cut into Yakima Basalt. 
Age--Pliocene (Hemphillian) . 
Collector- Shotwell, 1949. 
*Protosciurus condoni Black, 1963 
Type status-Holotype 
Specimen-F-5111. 
Locality- Exact locality not recorded. 
Stratigraphic position-] ohn Day Formation. 
Age- Late Oligocene or, more probably, early 
Miocene. 
Collector- Thomas Condon, 1870. 
*Citellus junturensis Shotwell, 1963 
Type status- Holotype 
Specimen-F-5811 , right lower jaw with P4-Ma. 
Locality-VO 2338. Malheur County, Oregon. 
Stratigraphic position-Juntura Formation, 
Upper Member, 50 ft. below pumice band. 
Age-Clarendonian, early Pliocene (late Miocene) 
Collector-Shotwell and Alvey, 1954. 
*Protosciurus rachelae Black 1963 
Type status-Hypotype 
Specimen-F-5039. 
Locality-Turtle Cove, exact horizon not recorded. 
Stratigraphic position-John Day Formation. 
Age-Probably early Miocene. 
Collector-Thomas Condon (?), 1869. 
*Leptodontomys oregonensis Shotwell, 1956 
Type status-Holotype 
Specimen-F-3633, right lower jaw with incisor 
and P4 • 
Locality-VO 2222, McKay Reservoir, east bank. 
Pendleton Quad. Umatilla County, Oregon. 
Stratigraphic position-Tuffaceous sandstone in 
channel cut into Yakima Basalt. 
Age- Pliocene (Hemphillian). 
Collector-]. A. Shotwell, 1956. 
*Adjidaumo quartzi Shotwell 1967 
Type status-Holotype 
Specimen-VO 22689, partial right mandible with 
P4- M2. 
Lccality-UO 2465, Quartz Basin, west side. 
Center of Sec. 33, T24S, R43E, WM, Malheur 
County, Oregon. 
Stratigraphic position-Deer Butte Formation. 
Age-Miocene (Barstovian). 
Collector-}. A. Shotwell, 1967. 
*Pseudotheridomys pagei Shotwell 1967 
Type status- Holotype 
6 BULLETIN, MUSEUM OF NATURAL HISTORY, UNIVERSITY OF OREGON No.24 
Specimen-VO 22715, mandible with P 4, M2-3 • 
Locality-VO 2465, Quartz Basin, west side; 
center of Sec. 33, T24S, R43E, WM, Malheur 
County, Oregon. 
Stratigraphic position-Deer Butte Formation. 
Age-Miocene (Barstovian). 
Collector- }. A. Shotwell, 1967. 
*Parapliosaccomys oregonensis Shotwell, 1967 
Type status-Holotype 
Specimen-VO 3631. 
Locality-VO 2222, McKay Reservoir, east bank; 
Pendleton Quad., Umatilla County, Oregon. 
Stratigraphic position-Channel cut into upper 
Yakima Basalt. 
Age-Pliocene ( Hemphillian) 
Collector-}. A. Shotwell, 1949. 
*Perognathus sargenti Shotwell, 1956 
Type status-Holotype 
Specimen-F-3637, left mandible with P 4-M3 • 
Locality-VO 2222, McKay Reservoir, east bank; 
Pendleton Quad., Umatilla County, Oregon. 
Stratigraphic position-Tuffaceous sandstone in 
channel cut into Yakima Basalt. 
Age-Pliocene (Hemphillian). 
Collector-}. A. Shotwell, 1949. 
*Dipoides smithi Shotwell, 1955 
Type status-Holotype 
Specimen-F -4072, left mandible with P 4-M2 
and incisor. 
Locality-VO 2222, McKay Reservoir, east bank; 
Pendleton Quad., Umatilla County, Oregon. 
Stratigraphic position-Tuffaceous sandstone in 
channel cut into upper Yakima Basalt. 
Age-Pliocene (Hemphillian). 
Collector-Shotwell, 1959. 
*Dipoides smithi Shotwell, 1955 
Type status-Paratypes 
Specimen-F-2820, palate with P 4-M2 of both 
sides; F-2821, lower jaw. 
Locality-U O 2222, McKay Reservoir, east bank; 
Pendleton Quad., Umatilla County, Oregon. 
Stratigraphic position-Same as holotype. 
Age-Pliocene (Hemphillian). 
Collector-Shotwell, 1949. 
*Eucastor malheurensis Shotwell, 1963 
Type status-Holotype 
Specimen-F-6683, right lower jaw with P 4-M2• 
Locality-VO 2352, Juniper Creek Canyon, west 
side, Malheur County, Oregon. 
Stratigraphu: position-Juntura Formation, 
Upper Member. 
Age-Clarendonian, early Pliocene (late 
Miocene). 
Collector-Shotwell. 
*Castor accessor Hay, 1927 (Shotwell, 1970). 
Type status-Plesiotype 
Specimen-VO 16338. 
Locality-VO 2404, unconsolidated, crossbedded 
sand lens; Grand View Fauna; Jackass Butte Quad., 
Idaho. 
Stratigraphic position-Glenns Ferry Formation. 
Age-late Pliocene. 
Collector-Shotwell, 1958. 
Comments-Shotwell ( 1970) considered Hay's 
type specimen of Castor accessor to be inadequate 
to characterize the species, and yet he did not want 
to create a new name for his specimens, which he 
considered to be conspecific with the C. accessor of 
Hay. He therefore states: '"I thus consider the new 
material as representing Castor accessor Hay and 
use it as a supplementary type in order that this 
beaver may be characterized." 
*Hystricops browni Shotwell, 1963 
Type status-Holotype 
Specimen-F-15696, associated partial dentition, 
with P4-M1 on each side. 
Locality-VO 2239, T21S, R35E; on highway 54 
east of Burns, Harney County, Oregon. 
Stratigraphic position-Drewsey Formation. 
Age-Pliocene (Hemphillian). 
Collector-Erikisson, 1952. 
*Procastoroides idahoensis Shotwell, 1970 
Type status-Holotype 
Specimen-VO 16267, lower left jaw with P 4-M2 • 
Locality-VO 2404, Jackass Butte Quad., Idaho. 
Stratigraphic position-Glenns Ferry Formation, 
in unconsolidated, cross-bedded sand lens; Grand 
View Fauna. 
Age-Blancan, late Pliocene. 
Collector-Shotwell, 1958. 
*Monosaulax typicus Shotwell, 1968 
Type status-Holotype 
Specimen-VO 21677. 
Locality-VO 2495, Skull Springs fauna, Red 
Basin; SE corner Sec. 10, T23S, R40E, WM, 
Malheur County, Oregon. 
Stratigraphic position-Butte Creek Volcanic 
Sandstone. 
Age-Miocene (Barstovian). 
Collector-Shotwell , 1961. 
*Dipoides vallicula Shotwell, 1970 
Type status-Holotype 
1983 GUST AF SON AND EATON: TYPE SPECIMENS 7 
Specimen-VO 26695, left mandible with P 4-M3. 
Locality-VO 2516, Little Valley, screening site 
#6406; Mitchell Butte Quad., Malheur County, 
Oregon. 
Stratigraphic position-Chalk Butte Formation. 
Age-Pliocene (Hemphillian). 
Collector-Shotwell and Russell , 1958. 
*Prosomys mimus Shotwell, 1956 
Promimomys mimus ( Shotwell 1956) Repenning, 
1968. 
Type status-Holotype 
Specimen-F-3667, right mandible with incisor 
and M1-2-
Locality-UO 2222, McKay Reservoir, east bank; 
Pendleton Quad., Umatilla County, Oregon. 
Stratigraphic position-Tuffaceous sandstone in 
channel cut into Yakima Basalt. 
Age-Pliocene (Hemphillian). 
Collector-Shotwell, 1949. 
*Peromyscus pagei Shotwell, 1967 
Type Status-Holotype 
Specimen-VO 22623, left mandible with M1-3. 
Locality-VO 2465, Quartz Basin, west side; 
center of Sec. 33, T24S, R42E, WM, Malheur County, 
Oregon. 
Stratigraphic position-Deer Butte Formation. 
Age-Miocene (Barstovian). 
Collector-Shotwell, 1960. 
*Peromyscus valensis Shotwell, 1967 
Type status-Holotype 
Specimen-VO 26920, mandible with incisor and 
M1-2-
Locality-UO 2516, Little Valley, screening site 
#6406, Mitchell Butte Quad., Malheur County, 
Oregon. 
Strat!·graphic position-Chalk Butte Formatio11. 
Age-Pliocene (Hemphillian). 
Collector-Shotwell and Russell, 1958. 
Order LAGOMORPHA 
*Ochotona spanglei Shotwell, 1956 
Type status-Holotype 
Specimen-F-4083, left lower jaw with P3- M3. 
Locality-VO 2222, McKay Reservoir, east bank; 
Pendleton Quad., Umatilla County, Oregon. 
Stratigraphic position-Tuffaceous sandstone in 
channel cut into Yakima Basalt. 
Age-Pliocene (Hemphillian) . 
Collector-Shotwell, 1949. 
*Hypolagus oregonensis Shotwell, 1956 
Type status-Holotype 
Specimen-F-4094, left lower jaw with P3-M3. 
Locality-VO 2222, McKay Reservoir, east hank; 
Pendleton Quad., Umatilla County, Oregon. 
Stratigraphic position-Tuffaceous sandstone in 
channel cut into Yakima Basalt. 
Age-Pliocene (Hemphillian). 
Collector-Shotwell, 1949. 
*Hypolagus oregonensis Shotwell, 1956 
Type status-Paratypes 
Specimen-F-4099, right maxilla fragment with 
P 2 ; F-2702, F-2907, F-4037, F-4036, F-2906, F-3064, 
F-2908, F-2063, F-3663, all lower jaws. 
Locality-VO 2222, McKay Reservoir, east bank; 
Pendleton Quad., Umatilla County, Oregon. 
Stratigraphic position-same as holotype. 
Age-Pliocene (Hemphillian). 
Collector-Shotwell, 1949. 
Order CARNIVORA 
*Canis condoni Shotwell, 1956 
Type status-Holotype 
Specimen-F-3241, lower jaw with M2. 
Locality-VO 2222, McKay Reservoir, Pendleton 
Quad., Umatilla County, Oregon. 
Stratigraphic position-Tuffaceous sandstone in 
channel cut into Yakima Basalt. 
Age- Pliocene (Hephillian). 
Collector-Shotwell, 1949. 
*Canis rurestris Condon, 1896 
Tomarctus rurestris (Condon, 1896) Downs, 1956 
Type Status-Holotype 
Specimen-VO 23077, skull and associated 
mandible. 
Locality-Cottonwood Creek, John Day Valley, 
Grant County, Oregon. 
Stratigraphic position-Mascall Formation, type 
locality. 
Age-Miocene (Hemingfordian, or possibly early 
Barstovian) . 
Collector- Thomas Condon, about 1870. 
*Sthenictus junturensis Shotwell, 1963 
Type Status-Holotype 
Specimen-F-6694, left lower jaw with P 2-M1• 
Locality-VO 2344, Malheur County, Oregon. 
Stratigraphic position-Juntura Formation, Upper 
Member. 
Age-Early Pliocene (Clarendonian). 
Collector-Shotwell and Alvey. 
*Martes (Plionictis) oregonensis Shotwell, 1970 
Type Status-Holotype 
" 
' 
8 BULLETIN, MUSEUM OF NATURAL HISTORY, UNIVERSITY OF OREGON No.24 
Specimen- F-26744, right mandible with P2--M2. 
Locality-UO 2516, Little Valley, 6406 screening 
site; Mitchell Butte Quad., Malheur County, Oregon. 
Stratigraphic position-Chalk Butte For'mation. 
Age- Pliocene (Hemphillian). 
Collector- Shotwell and Russell, 1958. 
*Felis longignathus Shotwell, 1956 
Type status- Holotype 
Specimen- F-2628, left lower jaw with C, PH, M1 • 
Locality-UO 2222, McKay Reservoir, Pendleton 
Quad., Umatilla County, Oregon. 
Stratigraphic position-Tuffaceous sandstone in 
channel in Yakima Basalt. 
Age-Pliocene (Hemphillian). 
Collector- Shotwell , 1949. 
*Desmatophoca oregonensis Condon, 1906 
Type status-Holotype 
Specimen-F-00735. 
Locality-UO 1153, Nye Beach, just west of 
Newport, Oregon. 
Stratigraphic position-Astoria Formation. 
Age-Middle Miocene. 
Collector- Thomas Condon, before 1906. 
Order PROBOSCIDEA 
*Mammut (Pliomastodon) furlongi Shotwell, 
1963 
Type Status-Holotype 
Specimen-F-10291, mandible with M1- 3 of both 
sides. 
Locality-UO 2448, Quarry 3, Malheur County, 
Oregon. 
Stratigraphic position- Juntura Formation, 
Upper Member. 
Age-Early Pliocene (Clarendonian). 
Collector-Shotwell, 1956. 
Order CETACEA 
*Cophocetus oregonensis Packard, 1934 
Type status-Holotype 
Specimen- F-00305, skull, parts of both forelimbs , 
vertebrae, innominates, and a few ribs. 
Locality-UO 1151, Newport, SE1;4, Sec. 7, TllS, 
Rll W, WM, Lincoln County, Oregon. 
Stratigraphic position-Astoria Formation. 
Age-Middle Miocene. 
Collector-Earl L. Packard, 1920. 
Order PERISSODACTYLA 
*Colodon (?) hancocki Radinsky, 1963 
Type Status-Holotype 
Specimen-UO 20377, partial skull. 
Locality-UO 2473, Clarno Mammal Quarry, 
Wheeler County, Oregon. 
Age-Early Oligocene ( early Chadronian) . 
Collector-Shotwell (?) 1960. 
*Hipparion condoni Merriam, 1915 
Type status-Holotype 
Specimen-#672 Condon Collection. 
Locality-Stone quarry at Ellensburg, 
Washington. 
Stratigraphic position-Ellensburg Formation. 
Age-Miocene. 
Collector-Sent to Condon by W.R. Abrams 
before 1902. 
Order ARTIODACTYLA 
*Prosthennops brachirostris Shotwell, 1956 
Type Status-Holotype 
Specimen-F-2767, palate with canines and all 
cheek teeth. 
Locality-UO 2222, McKay Reservoir, Pendleton 
Quad., Umatilla County, Oregon. 
Stratigraphic position-Tuffaceous sandstone in 
channel cut into Yakima Basalt. 
Age-Pliocene (Hemphillian). 
Collector-Shotwell, 1949. 
*Prosthennops brachirostris Shotwell, 1956 
Type status-Paratype 
Specimen-F-3585, lower jaw with canines, 
P4-M3. 
Locality-UO 2222, McKay Reservoir, east bank, 
Pendleton Quad., Umatilla County, Oregon. 
Stratigraphic position-Tuffaceous sandstone in 
channel cut into Yakima Basalt. 
Age-Pliocene (Hemphillian). 
Collector-Shotwell, 1949. 
*Dicotyles pristinus Leidy, 1873 
Perchoerus pristinus (Leidy, 1873) Cope and 
Matthew, 1915 
Type status-Holotype 
Specimen-F-5049, skull. 
Locality-UO NFJD, North Fork of John Day 
River, eastern Oregon. 
Stratigraphic position-] ohn Day Formation. 
Age-Miocene. 
Collector-Thomas Condon. 
*Palaeochoerus subaequans Cope, 1879 
Perchoerus subaequans (Cope, 1879) Cope and 
Matthew, 1915 
Type status-Holotype 
Specimen- F-669. 
1983 GUSTAFSON AND EATON : TYPE SPECIMENS 9 
Locality-VO NFJD, North Fork of John Day 
River, eastern Oregon. 
Stratigraphic position-] ohn Day Formation. 
Age-Miocene (Late Arikareean). 
Collector- Thomas Condon. 
*Chaenohyos decedens Cope, 1879 
Perchoerus decedens (Cope, 1879) Cope and 
Matthew, 1915. 
Type status-Holotype 
Specimen- F-669. 
Locality-VO NFJD, North Fork of John Day 
River, eastern Oregon. 
Stratigraphic position-John Day Formation. 
Age- Miocene (Late Arikareean). 
Collector- Thomas Condon. 
*Oreodon soperbos Leidy, 1870 
Promerycochoerus superbus (Leidy, 1870) 
Douglass, 1901 
Type status-Holotype 
Specimen-F-5146, F-5147, F-5148, F-5342, 
F-5361, mutilated skull (Condon yellow numbers 
718, 721, 722, 723, 726, and 727). 
Locality-Bridge Creek, John Day Basin, Oregon. 
Stratigraphic position-John Day Formation. 
Age-Miocene (Late Arikareean). 
Collector-Thomas Condon, about 1869. 
Comments-For many years this specimen was 
thought to be lost (see Schultz and Falkenbach, 1949: 
107-108). It was reidentified by direct comparison 
with Leidy's illustration of 1873. Comparison was 
difficult because the specimen is in several pieces, 
and the illustrator had drawn the snout from the op-
posite side of the specimen. Leidy's illustration is re-
produced on the cover of this Bulletin. 
*Oreodon soperbos Leidy, 1870 
Promerycochoerus superbus (Leidy, 1870) 
Douglass, 1901 
Type status- Cotype 
Specimen-F-684. 
Locality-Turtle Cove, John Day Basin, Oregon. 
Stratigraphic position-John Day Formation. 
Age-Miocene (Late Arikareean). 
Collector-Thomas Condon. 
PLANTS 
Family Awacardiaceae 
*Rhos varians Lakhanpal, 1958 
Type status-Syntype 
Specimen-F-4290, 5 leaflets around stem. 
Locality-VO 2715, Willamette Flora, 14 mile 
north of Goshen, Lane County, Oregon. 
Stratigraphic position-? Eugene Formation 
( tuffaceous shales and sandstone, which may overlie 
Eugene Fm., Fisher Fm., or both). 
Age-Oligocene. 
Collector -Chaney. 
*Rhos varians Lakhanpal, 1958 
Type status-Syntype 
Specimen- F-4291, single leaflet on slab. 
Locality-VO 2715, Willamette Flora (as above). 
Stratigraphic position- ? Eugene Formation (see 
above). 
Age-Oligocene. 
Collector-Chaney. 
*Rhos varians Lakhanpal, 1958 
Type status-Paratype 
Specimen-F-4202. 
Locality-VO 2715, Willamette Flora (as above). 
Stratigraphic position- ? Eugene Formation (see 
above) . 
Age-Oligocene. 
Collector-Chaney. 
10 BULLETIN, MUSEUM OF NATURAL HISTORY , UNIVERSITY OF OREGON No.24 
LITERATURE CITED 
Black, C. C., 1963. A review of the North American 
Tertiary Sciuridae. Bull. of the Museum of Com-
parative Zoology, Harvard Univ., 103(3): 109-
248. 
Brodkorb, Pierce, 1958. Birds from the Middle 
Pliocene of McKay, Oregon. The Condor, 60 : 252-
255. 
, 1961. Birds from the Pliocen.e of Jun-
tura, Oregon. Quart. }our. Florida Acad. Sci., 
24(3) :169-184. 
Cavender, T.M., and R.R. Miller, 1972. Smilodo-
nichthyes rastrosus: a new Pliocene salmonid fish. 
Bull . Univ. of Oregon Museum of Natural History, 
18:1-44. 
Condon, Thomas, 1896. Scientific description of 
two new fossil dogs. Bull. Univ. Oregon, 2 ( 6) : 
1-11. 
-----, 1906. A new fossil pinniped (Desmato-
phoca oregonensis). Bull. Univ. Oregon, 3 ( 33) : 
1-14. 
Cope, E. D., 1879. Second contribution to a knowl-
edge of the Miocene fauna of Oregon. Proc. Ameri-
can Philos. Soc., 18:370-376. 
-----, and Matthew, W. D., 1915. Tertiary 
Mammalia and Permian Vertebrata. American 
Museum of Natural History, Monograph series no. 
2, 68 plates including explanations. 
Douglass, Earl, 1901. New species of Merycochoe-
rus in Montana, part II. American Jour. of Sci. 
ser. 4, v. 11 :73-83. 
Downs, T., 1956. The Mascall fauna from the Mio-
cene of Oregon. Univ. California Puhl. in Geol. 
Sci., 31 ( 5) : 199-354. 
Hay, 0. P., 1927. The Pleistocene of the western 
region of North America and its vertebrated ani-
mals. Publications of the Carnegie Institution of 
Washington, 3228, 346 pp. 
Hutchison, J. H., 1966. Notes ()n some upper Mio-
cene shrews from Oregon. Bull. Univ. of Oregon 
Museum of Natural History, 2:1-23. 
-----, 1968. Fossil Talpidae (lnsectivora, 
Mammalia) from the later Tertiary of Oregon. 
Bull. Univ. of Oregon Museum of Natural History, 
11:1-117. 
Lakhanpal, R. N. 1958. The Rujada Flora of west 
central Oregon. Univ. California Puhl. in Grol. Sci. , 
35 ( 1) :l-66. 
Langston, W., C. C. Black, J. A. Lillegraven, 
T. H. Patton, B. Schaeffer, and R. W. Wilson, 
1977. Fossil vertebrates in the United States, the 
next ten years. A report of the Society of Verte-
brate Paleontology Advisory Committee for Sys-
tematic Resources in Vertebrate Paleontology to 
the National Science Foundati()n. Soc. Vert. Paleo., 
Dec.1977, 44p. 
Leidy, Joseph, 1870. Remarks on a collection of 
fossils from Dalles City, Oregon. Proc. Acad. Nat. 
Sci. Philadelphia, 1870, 111-113. 
-----, 1873. Contributions to the extinct ver-
tebrate fauna of the western territories. Part I of 
volume I, Fossil Vertebrates. Report t() the U.S. 
Geol. Surv. of the Territories (F. V. Hayden, Geol-
ogist in charge). Washington, D.C., 1-358, 37 
plates. 
Lundberg, J. G., 1975. The fossil catfish of North 
America. Museum of Paleontology, Univ. of Mich-
igan; Papers on Paleontolgy No. 11; Claude W. 
Hibbard Memorial Vol. 2:1-51. 
Merriam, J.C., 1915. New species of the Hipparion 
group from the Pacific Coast and Great Basin prov-
inces of North America. Univ. of California Pub-
lications, Bull. in Geology, 9 ( 1) : 1-8. 
Packard, E. L., and R. Kellogg, 1934. A new ce-
tothere from the Miocene Astoria Formation of 
Newport, Oregon. Contributions to Paleontol()gy: 
Marine Mammals, Carnegie Institution of Wash-
ington, Puhl. 44 7: 1-62. 
Radinsky, Leonard, 1963. Origin and early evolu-
tion of North American Tapiroidea. Bull. Peabody 
Museum of Natural History, 17:1-106. 
Repenning, C. A., 1967. Subfamilies and genera 
of the Soricidae. U.S. Ge-01. Surv. Prof. Pap. 565: 
1-74. 
-----, 1968. Mandibular musculature and the 
ongm of the subfamily Arvicolinae (Rodentia). 
Acta Zoologica Cracoviensia, 8 :29-72. 
-----, and R. H. Tedford, 1977. Otarioid seals 
of the Neogene. U. S. Geol. Surv. Prof. Pap. 992: 
1-93, 24 plates. 
1983 GUST AF SON AND EATON: TYPE SPECIMENS 11 
Schultz, C. B., and Falkenbach, C. H., 1949. Pro-
merycochoerinae, a new subfamily of oreodonts. 
Bull. of the American Museum of Natural History 
93(3) :71-198. ' 
Shotwell, J. A., 1955. Review of the Pliocene beaver 
Dipoides. Jour. of Paleontology, 29(1) :129-144. 
-----, 1956. Hemphillian mammalian assem-
blage from northeastern Oregon. Bull. Geo!. Soc. 
America, 67: 717 -738. 
-----, 1?58. Evolution and biogeography of 
the aplodontid and mylagaulid rodents. Evolution. 
12 ( 4) :451-484. -
----- , 1963. The Juntura Basin: Studies in 
earth history and paleoecology. Trans. American 
Phil. Soc., 53 ( 1) : 1-77. 
-----, 1967. Peromyscus of the late Tertiary in 
Oregon. Bull. Univ. of Oregon Museum of Natural 
History, 5: 1-35. 
. , 1967. Late Tertiary geomyoid rodents 
m Oregon. Bull. Univ. of Oregon Museum of Nat-
ural History, 9:1-51. 
-----, 1968. Miocene mammals of southeast 
Oregon. Bull. Univ. of Oregon Museum of Natural 
History, 14:1-67. 
------, 1970. Pliocene mammals of southeast 
Oregon and adjacent Idaho. Bull. Univ. of Oregon 
Museum of Natural History, 17:1-103. 
van Frank, R. V., 1955. Paleotaricha oligocenica 
new genus and species, an Oligocene salamande; 
from Oregon. Breviora, Museum of Comparative 
Zoology, 45:1-15. 

f continued from inside front cover .1 
BuUetins 
No. 11 Fossil Talpidae (lnsectivora, Mammalia) from the Tertiary of Oregon, by J. H. Hutchi-
son ; 117 pages, 98 figures, July 1968 
No. 12 Plants of the Three Sisters Region, Oregon Cascade Range, by Orlin L. Ireland; 130 
pages, 34 figures, April 1968 
No. 13 Historical Background of the Flora of the Pacific Northwest, by LeRoy Detling; 57 pages, 
6 figures , July 1968 
No. 14 Miocene Mammals of Southeast Oregon, by J. Arnold Shotwell ; 67 pages, 33 figures , 
Price 
$1.25 
$3.75 
$1.50 
August 1968 $1.25 
No. 15 Item Seriation as an Aid for Elementary Scale and Cluster Analysis, by LeRoy Johnson, 
Jr.; 46 pages, 19 figures, September 1968 
No. 16 The Oligocene Marine Molluscan Fauna of the Eugene Formation in Oregon, by Carole 
Jean Stentz Hickman; 11 pages, 14 plates, 4 figures, August 1969 
No.17 Pliocene Mammals of Southeast Oregon and Adjacent Idaho, by J. Arnold Shotwell; 103 
pages, 42 figures, August 1970 
No. 18 Smilodonichthys rastrosus, a New Pliocene Salmonid Fish from Western United States, 
by TedM. Cavender and Robert Rush Miller; 44 pages, 14 figures, March 1972 
No. 19 Journal of First Trip of University of California to John Day Beds of Eastern Oregon, by 
Loye Miller, edited by J. Arnold Shotwell; 21 pages, 7 figures, 1 map, December 1971, sec-
ond printing April 1983 
No. 20 An Approach to the Study of Far Western North American Prehistory: Early Man, by 
Luther S. Cressman; 11 pages, August 1973 
No. 21 Guide to the Geology of the Owyhee Region of Oregon, by Laurence R. Kittleman; 61 
pages, 4 plates, 36 figures, 3 maps, September 1973 
No. 22 Guide to the Geology and Lore of the Wild Reach of the Rogue River, Oregon, by William 
B. Purdom; 67 pages, 4 plates, 31 figures, 7 maps, May 1977 
No. 23 The Vertebrate Faunas of the Pliocene Ringld Formation, South-Central Washington, by 
Eric Paul Gustafson; 62 pages, 31 figures, March 1978 
No. 24 Type Specimens at the University of Oregon: Fossil Vertebrates and Plants, by Eric Paul 
$1.50 
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Gustafson and Joyce Eaton; 11 pages, March 1983 $2.00 
;